SigPhi · Charles Darwin

The Variation of Animals and Plants under Domestication

Page 16 of 72

rupestris_ of Asia. Now if we turn to the domestic races, it is highly remarkable, as an eminent fancier, Mr. Wicking, observed to me, that, whenever a blue bird appears in any race, the wings almost invariably show the double black bars.[23] The primary wing-feathers may be white or black, and the whole body may be of any colour, but if the wing-coverts are blue, the two black bars are sure to appear. I have myself seen, or acquired trustworthy evidence, as given below,[24] of blue birds with black bars on the wing, with the croup either white or very pale or dark blue, with the tail having a terminal black bar, and with the outer feathers externally edged with white or very pale coloured, in the following races, which, as I carefully observed in each case, appeared to be perfectly true: namely, in Pouters, Fantails, Tumblers, Jacobins, Turbits, Barbs, Carriers, Runts of three distinct varieties, Trumpeters, Swallows, and in many other toy-pigeons, which as being closely allied to _C. livia,_ are not worth enumerating. Thus we see that, in purely-bred races of every kind known in Europe, blue birds occasionally appear having all the marks which characterise _C. livia,_ and which concur in no other wild species. Mr. Blyth, also, has made the same observation with respect to the various domestic races known in India.

Certain variations in the plumage are equally common in the wild _C. livia,_ in dovecot-pigeons, and in all the most highly modified races. Thus, in all, the croup varies from white to blue, being most frequently white in Europe, and very generally blue in India.[25] We have seen that the wild _C. livia_ in Europe, and dovecots in all parts of the world, often have the upper wing-coverts chequered with black; and all the most distinct races, when blue, are occasionally chequered in precisely the same manner.

Thus I have seen Pouters, Fantails, Carriers, Turbits, Tumblers (Indian and English), Swallows, Bald-pates, and other toy-pigeons blue and chequered; and Mr. Esquilant has seen a chequered Runt. I bred from two pure blue Tumblers a chequered bird.

The facts hitherto given refer to the occasional appearance in pure races of blue birds with black wing-bars, and likewise of blue and chequered birds; but it will now be seen that when two birds belonging to distinct races are crossed, neither of which have, nor probably have had during many generations, a trace of blue in their plumage, or a trace of wing-bars and the other characteristic marks, they very frequently produce mongrel offspring of a blue colour, sometimes chequered, with black wing-bars, etc.; or if not of a blue colour, yet with the several characteristic marks more or less plainly developed. I was led to investigate this subject from MM. Boitard and Corbié[26] having asserted that from crosses between certain breeds it is rare to get anything but bisets or dovecot pigeons, which, as we know, are blue birds with the usual characteristic marks. We shall hereafter see that this subject possesses, independently of our present object, considerable interest, so that I will give the results of my own trials in full.

I selected for experiment races which, when pure, very seldom produce birds of a blue colour, or have bars on their wings and tail.

The Nun is white, with the head, tail, and primary wing-feathers black; it is a breed which was established as long ago as the year 1600. I crossed a male Nun with a female red common Tumbler, which latter variety generally breeds true. Thus neither parent had a trace of blue in the plumage, or of bars on the wing and tail. I should premise that common Tumblers are rarely blue in England. From the above cross I reared several young: one was red over the whole back, but with the tail as blue as that of the rock-pigeon; the terminal bar, however, was absent, but the outer feathers were edged with white: a second and third nearly resembled the first, but the tail in both presented a trace of the bar at the end: a fourth was brownish, and the wings showed a trace of the double bar: a fifth was pale blue over the whole breast, back, croup, and tail, but the neck and primary wing-feathers were reddish; the wings presented two distinct bars of a red colour; the tail was not barred, but the outer feathers were edged with white.

I crossed this last curiously coloured bird with a black mongrel of complicated descent, namely, from a black Barb, a Spot, and Almond-tumbler, so that the two young birds produced from this cross included the blood of five varieties, none of which had a trace of blue or of wing and tail-bars: one of the two young birds was brownish-black, with black wing-bars; the other was reddish-dun, with reddish wing-bars, paler than the rest of the body, with the croup pale blue, the tail bluish with a trace of the terminal bar.

Mr. Eaton[27] matched two Short-faced Tumblers, namely, a splash cock and kite hen (neither of which are blue or barred), and from the first nest he got a perfect blue bird, and from the second a silver or pale blue bird, both of which, in accordance with all analogy, no doubt presented the usual characteristic marks.

I crossed two male black Barbs with two female red Spots. These latter have the whole body and wings white, with a spot on the forehead, the tail and tail-coverts red; the race existed at least as long ago as 1676, and now breeds perfectly true, as was known to be the case in the year 1735.[28] Barbs are uniformly-coloured birds, with rarely even a trace of bars on the wing or tail; they are known to breed very true. The mongrels thus raised were black or nearly black, or dark or pale brown, sometimes slightly piebald with white: of these birds no less than six presented double wing-bars; in two the bars were conspicuous and quite black; in seven some white feathers appeared on the croup; and in two or three there was a trace of the terminal bar to the tail, but in none were the outer tail-feathers edged with white.

I crossed black Barbs (of two excellent strains) with purely-bred, snow-white Fantails. The mongrels were generally quite black, with a few of the primary wing and tail feathers white: others were dark reddish-brown, and others snow-white: none had a trace of wing-bars or of the white croup. I then paired together two of these mongrels, namely, a brown and black bird, and their offspring displayed wing-bars, faint, but of a darker brown than the rest of body. In a second brood from the same parents a brown bird was produced, with several white feathers confined to the croup.

I crossed a male dun Dragon belonging to a family which had been dun- coloured without wing-bars during several generations, with a uniform red Barb (bred from two black Barbs); and the offspring presented decided but faint traces of wing-bars. I crossed a uniform red male Runt with a White trumpeter; and the offspring had a slaty-blue tail with a bar at the end, and with the outer feathers edged with white. I also crossed a female black and white chequered Trumpeter (of a different strain from the last) with a male Almond-tumbler, neither of which exhibited a trace of blue, or of the white croup, or of the bar at end of tail: nor is it probable that the progenitors of these two birds had for many generations exhibited any of these characters, for I have never even heard of a blue Trumpeter in this country, and my Almond-tumbler was purely bred; yet the tail of this mongrel was bluish, with a broad black bar at the end, and the croup was perfectly white. It may be observed in several of these cases, that the tail first shows a tendency to become by reversion blue; and this fact of the persistency of colour in the tail and tail-coverts[29] will surprise no one who has attended to the crossing of pigeons.

The last case which I will give is the most curious. I paired a mongrel female Barb-fantail with a mongrel male Barb-spot; neither of which mongrels had the least blue about them. Let it be remembered that blue Barbs are excessively rare; that Spots, as has been already stated, were perfectly characterised in the year 1676, and breed perfectly true; this likewise is the case with white Fantails, so much so that I have never heard of white Fantails throwing any other colour.

Nevertheless the offspring from the above two mongrels was of exactly the same blue tint as that of the wild rock-pigeon from the Shetland Islands over the whole back and wings; the double black wing-bars were equally conspicuous; the tail was exactly alike in all its characters, and the croup was pure white; the head, however, was tinted with a shade of red, evidently derived from the Spot, and was of a paler blue than in the rock-pigeon, as was the stomach. So that two black Barbs, a red Spot, and a white Fantail, as the four purely-bred grandparents, produced a bird exhibiting the general blue colour, together with every characteristic mark, the wild _Columba livia._ With respect to crossed breeds frequently producing blue birds chequered with black, and resembling in all respects both the dovecot-pigeon and the chequered wild variety of the rock-pigeon, the statement before referred to by MM. Boitard and Corbié would almost suffice; but I will give three instances of the appearance of such birds from crosses in which one alone of the parents or great-grandparents was blue, but not chequered. I crossed a male blue Turbit with a snow-white Trumpeter, and the following year with a dark, leaden-brown, Short-faced Tumbler; the offspring from the first cross were as perfectly chequered as any dovecot-pigeon; and from the second, so much so as to be nearly as black as the most darkly chequered rock-pigeon from Madeira. Another bird, whose great-grandparents were a white Trumpeter, a white Fantail, a white Red-spot, a red Runt, and a blue Pouter, was slaty-blue and chequered exactly like a dovecot-pigeon. I may here add a remark made to me by Mr. Wicking, who has had more experience than any other person in England in breeding pigeons of various colours: namely, that when a blue, or a blue and chequered bird, having black wing- bars, once appears in any race and is allowed to breed, these characters are so strongly transmitted that it is extremely difficult to eradicate them.

What, then, are we to conclude from this tendency in all the chief domestic races, both when purely bred and more especially when intercrossed, to produce offspring of a blue colour, with the same characteristic marks, varying in the same manner, as in _Columbia livia_? If we admit that these races are all descended from _C. livia,_ no breeder will doubt that the occasional appearance of blue birds thus characterised is accounted for on the well-known principle of “throwing back” or reversion. Why crossing should give so strong a tendency to reversion, we do not with certainty know; but abundant evidence of this fact will be given in the following chapters. It is probable that I might have bred even for a century pure black Barbs, Spots, Nuns, white Fantails, Trumpeters, etc., without obtaining a single blue or barred bird; yet by crossing these breeds I reared in the first and second generation, during the course of only three or four years, a considerable number of young birds, more or less plainly coloured blue, and with most of the characteristic marks. When black and white, or black and red birds, are crossed, it would appear that a slight tendency exists in both parents to produce blue offspring, and that this, when combined, overpowers the separate tendency in either parent to produce black, or white, or red offspring.

If we reject the belief that all the races of the pigeon are the modified descendants of _C. livia,_ and suppose that they are descended from several aboriginal stocks, then we must choose between the three following assumptions: firstly, that at least eight or nine species formerly existed which were aboriginally coloured in various ways, but have since varied in exactly the same manner so as to assume the colouring of _C. livia_; but this assumption throws not the least light on the appearance of such colours and marks when the races are crossed.

Or secondly, we may assume that the aboriginal species were all coloured blue, and had the wing-bars and other characteristic marks of _C. livia,_—a supposition which is highly improbable, as besides this one species no existing member of the Columbidæ presents these combined characters; and it would not be possible to find any other instance of several species identical in plumage, yet as different in important points of structure as are Pouters, Fantails, Carriers, Tumblers, etc.

Or lastly, we may assume that all the races, whether descended from _C.

livia_ or from several aboriginal species, although they have been bred with so much care and are so highly valued by fanciers, have all been crossed within a dozen or score of generations with _C. livia,_ and have thus acquired their tendency to produce blue birds with the several characteristic marks. I have said that it must be assumed that each race has been crossed with _C. livia_ within a dozen, or, at the utmost, within a score of generations; for there is no reason to believe that crossed offspring ever revert to one of their ancestors when removed by a greater number of generations. In a breed which has been crossed only once, the tendency to reversion will naturally become less and less in the succeeding generations, as in each there will be less and less of the blood of the foreign breed; but when there has been no cross with a distinct breed, and there is a tendency in both parents to revert to some long-lost character, this tendency, for all that we can see to the contrary, may be transmitted undiminished for an indefinite number of generations. These two distinct cases of reversion are often confounded together by those who have written on inheritance.

Considering, on the one hand, the improbability of the three assumptions which have just been discussed, and, on the other hand, how simply the facts are explained on the principle of reversion, we may conclude that the occasional appearance in all the races, both when purely bred and more especially when crossed, of blue birds, sometimes chequered, with double wing-bars, with white or blue croups, with a bar at the end of the tail, and with the outer tail-feathers edged with white, affords an argument of the greatest weight in favour of the view that all are descended from _Columba livia,_ including under this name the three or four wild varieties or sub-species before enumerated.

To sum up the six foregoing arguments, which are opposed to the belief that the chief domestic races are the descendants of at least eight or nine or perhaps a dozen species; for the crossing of any less number would not yield the characteristic differences between the several races. _ Firstly,_ the improbability that so many species should still exist somewhere, but be unknown to ornithologists, or that they should have become within the historical period extinct, although man has had so little influence in exterminating the wild _C. livia. Secondly,_ the improbability of man in former times having thoroughly domesticated and rendered fertile under confinement so many species. _Thirdly,_ these supposed species having nowhere become feral. _Fourthly,_ the extraordinary fact that man should, intentionally or by chance, have chosen for domestication several species, extremely abnormal in character; and furthermore, the points of structure which render these supposed species so abnormal being now highly variable. _Fifthly,_ the fact of all the races, though differing in many important points of structure, producing perfectly fertile mongrels; whilst all the hybrids which have been produced between even closely allied species in the pigeon-family are sterile. _Sixthly,_ the remarkable statements just given on the tendency in all the races, both when purely bred and when crossed, to revert in numerous minute details of colouring to the character of the wild rock-pigeon, and to vary in a similar manner. To these arguments may be added the extreme improbability that a number of species formerly existed, which differed greatly from each other in some few points, but which resembled each other as closely as do the domestic races in other points of structure, in voice, and in all their habits of life. When these several facts and arguments are fairly taken into consideration, it would require an overwhelming amount of evidence to make us admit that the chief domestic races are descended from several aboriginal stocks; and of such evidence there is absolutely none.

The belief that the chief domestic races are descended from several wild stocks no doubt has arisen from the apparent improbability of such great modifications of structure having been effected since man first domesticated the rock-pigeon. Nor am I surprised at any degree of hesitation in admitting their common parentage: formerly, when I went into my aviaries and watched such birds as Pouters, Carriers, Barbs, Fantails, and Short-faced Tumblers, etc., I could not persuade myself that all had descended from the same wild stock, and that man had consequently in one sense created these remarkable modifications.

Therefore I have argued the question of their origin at great, and, as some will think, superfluous length.

Finally, in favour of the belief that all the races are descended from a single stock, we have in _Columba livia_ a still existing and widely distributed species, which can be and has been domesticated in various countries. This species agrees in most points of structure and in all its habits of life, as well as occasionally in every detail of plumage, with the several domestic races. It breeds freely with them, and produces fertile offspring.

It varies in a state of nature,[30] and still more so when semi-domesticated, as shown by comparing the Sierra Leone pigeons with those of India, or with those which apparently have run wild in Madeira. It has undergone a still greater amount of variation in the case of the numerous toy-pigeons, which no one supposes to be descended from distinct species; yet some of these toy-pigeons have transmitted their character truly for centuries. Why, then, should we hesitate to believe in that greater amount of variation which is necessary for the production of the eleven chief races? It should be borne in mind that in two of the most strongly-marked races, namely, Carriers and Short-faced Tumblers, the extreme forms can be connected with the parent-species by graduated differences not greater than those which may be observed between the dovecot-pigeons inhabiting different countries, or between the various kinds of toy-pigeons,—gradations which must certainly be attributed to variation.

That circumstances have been eminently favourable for the modification of the pigeon through variation and selection will now be shown. The earliest record, as has been pointed out to me by Professor Lepsius, of pigeons in a domesticated condition, occurs in the fifth Egyptian dynasty, about 3000 B.C.;[31] but Mr. Birch, of the British Museum, informs me that the pigeon appears in a bill of fare in the previous dynasty. Domestic pigeons are mentioned in Genesis, Leviticus, and Isaiah.[32] In the time of the Romans, as we hear from Pliny,[33] immense prices were given for pigeons; “nay, they are come to this pass, that they can reckon up their pedigree and race.” In India, about the year 1600, pigeons were much valued by Akbar Khan: 20,000 birds were carried about with the court, and the merchants brought valuable collections. “The monarch of Iran and Turan sent him some very rare breeds. His Majesty,” says the courtly historian, “by crossing the breeds, which method was never practised before, has improved them astonishingly.”[34] Akber Khan possessed seventeen distinct kinds, eight of which were valuable for beauty alone. At about this same period of 1600 the Dutch, according to Aldrovandi, were as eager about pigeons as the Romans had formerly been. The breeds which were kept during the fifteenth century in Europe and in India apparently differed from each other. Tavernier, in his Travels in 1677, speaks, as does Chardin in 1735, of the vast number of pigeon-houses in Persia; and the former remarks that, as Christians were not permitted to keep pigeons, some of the vulgar actually turned Mahometans for this sole purpose. The Emperor of Morocco had his favourite keeper of pigeons, as is mentioned in Moore’s treatise, published 1737. In England, from the time of Willughby in 1678 to the present day, as well as in Germany and in France, numerous treatises have been published on the pigeon. In India, about a hundred years ago, a Persian treatise was written; and the writer thought it no light affair, for he begins with a solemn invocation, “in the name of God, the gracious and merciful.” Many large towns, in Europe and the United States, now have their societies of devoted pigeon-fanciers: at present there are three such societies in London. In India, as I hear from Mr. Blyth, the inhabitants of Delhi and of some other great cities are eager fanciers. Mr. Layard informs me that most of the known breeds are kept in Ceylon. In China, according to Mr. Swinhoe of Amoy, and Dr. Lockhart of Shangai, Carriers, Fantails, Tumblers, and other varieties are reared with care, especially by the bonzes or priests. The Chinese fasten a kind of whistle to the tail-feathers of their pigeons, and as the flock wheels through the air they produce a sweet sound. In Egypt the late Abbas Pacha was a great fancier of Fantails. Many pigeons are kept at Cairo and Constantinople, and these have lately been imported by native merchants, as I hear from Sir W. Elliot, into Southern India, and sold at high prices.

The foregoing statements show in how many countries, and during how long a period, many men have been passionately devoted to the breeding of pigeons. Hear how an enthusiastic fancier at the present day writes: “If it were possible for noblemen and gentlemen to know the amazing amount of solace and pleasure derived from Almond Tumblers, when they begin to understand their properties, I should think that scarce any nobleman or gentleman would be without their aviaries of Almond Tumblers.”[35] The pleasure thus taken is of paramount importance, as it leads amateurs carefully to note and preserve each slight deviation of structure which strikes their fancy. Pigeons are often closely confined during their whole lives; they do not partake of their naturally varied diet; they have often been transported from one climate to another; and all these changes in their conditions of life would be likely to cause variability.

Pigeons have been domesticated for nearly 5000 years, and have been kept in many places, so that the numbers reared under domestication must have been enormous: and this is another circumstance of high importance, for it obviously favours the chance of rare modifications of structure occasionally appearing. Slight variations of all kinds would almost certainly be observed, and, if valued, would, owing to the following circumstances, be preserved and propagated with unusual facility. Pigeons, differently from any other domesticated animal, can easily be mated for life, and, though kept with other pigeons, rarely prove unfaithful to each other. Even when the male does break his marriage-vow, he does not permanently desert his mate. I have bred in the same aviaries many pigeons of different kinds, and never reared a single bird of an impure strain. Hence a fancier can with the greatest ease select and match his birds. He will also see the good results of his care; for pigeons breed with extraordinary rapidity. He may freely reject inferior birds, as they serve at an early age as excellent food.

_History of the principal Races of the Pigeon._[36] Before discussing the means and steps by which the chief races have been formed, it will be advisable to give some historical details, for more is known of the history of the pigeon, little though this is, than of any other domesticated animal. Some of the cases are interesting as proving how long domestic varieties may be propagated with exactly the same or nearly the same characters; and other cases are still more interesting as showing how slowly but steadily races have been greatly modified during successive generations. In the last chapter I stated that Trumpeters and Laughers, both so remarkable for their voices, seem to have been perfectly characterised in 1735; and Laughers were apparently known in India before the year 1600. Spots in 1676, and Nuns in the time of Aldrovandi, before 1600, were coloured exactly as they now are. Common Tumblers and Ground Tumblers displayed in India, before the year 1600, the same extraordinary peculiarities of flight as at the present day, for they are well described in the ‘Ayeen Akbery.’ These breeds may all have existed for a much longer period; we know only that they were perfectly characterised at the dates above given. The _ average_ length of life of the domestic pigeon is probably about five or six years; if so, some of these races have retained their character perfectly for at least forty or fifty generations.

_Pouters._—These birds, as far as a very short description serves for comparison, appear to have been well characterised in Aldrovandi’s time,[37] before the year 1600. Length of body and length of leg are at the present time the two chief points of excellence. In 1735 Moore said (see Mr. J. M. Eaton’s edition)—and Moore was a first-rate fancier—that he once saw a bird with a body 20 inches in length, “though 17 or 18 inches is reckoned a very good length;” and he has seen the legs very nearly 7 inches in length, yet a leg 6½ or 6¾ long “must be allowed to be a very good one.” Mr. Bult, the most successful breeder of Pouters in the world, informs me that at present (1858) the standard length of the body is not less than 18 inches; but he has measured one bird 19 inches in length, and has heard of 20 and 22 inches, but doubts the truth of these latter statements. The standard length of the leg is now 7 inches, but Mr. Bult has recently measured two of his own birds with legs 7½ long. So that in the 123 years which have elapsed since 1735 there has been hardly any increase in the standard length of the body; 17 or 18 inches was formerly reckoned a very good length, and now 18 inches is the minimum standard; but the length of leg seems to have increased, as Moore never saw one quite 7 inches long; now the standard is 7, and two of Mr. Bult’s birds measured 7½ inches in length. The extremely slight improvement in Pouters, except in the length of the leg, during the last 123 years, may be partly accounted for by the neglect which they suffered, as I am informed by Mr. Bult, until within the last 20 or 30 years. About 1765[38] there was a change of fashion, stouter and more feathered legs being preferred to thin and nearly naked legs.

_Fantails._—The first notice of the existence of this breed is in India, before the year 1600, as given in the ‘Ayeen Akbery;’[39] at this date, judging from Aldrovandi, the breed was unknown in Europe. In 1677 Willughby speaks of a Fantail with 26 tail-feathers; in 1735 Moore saw one with 36 feathers; and in 1824 MM. Boitard and Corbié assert that in France birds can easily be found with 42 tail-feathers. In England, the number of the tail-feathers is not at present so much regarded as their upward direction and expansion. The general carriage of the bird is likewise now much valued. The old descriptions do not suffice to show whether in these latter respects there has been much improvement: but if Fantails with their heads and tails touching had formerly existed, as at the present time, the fact would almost certainly have been noticed. The Fantails which are now found in India probably show the state of the race, as far as carriage is concerned, at the date of their introduction into Europe; and some, said to have been brought from Calcutta, which I kept alive, were in a marked manner inferior to our exhibition birds. The Java Fantail shows the same difference in carriage; and although Mr.

Swinhoe has counted 18 and 24 tail-feathers in his birds, a first-rate specimen sent to me had only 14 tail-feathers.

_Jacobins._—This breed existed before 1600, but the hood, judging from the figure given by Aldrovandi, did not enclose the head nearly so perfectly as at present: nor was the head then white; nor were the wings and tail so long, but this last character might have been overlooked by the rude artist. In Moore’s time, in 1735, the Jacobin was considered the smallest kind of pigeon, and the bill is said to be very short. Hence either the Jacobin, or the other kinds with which it was then compared, must since that time have been considerably modified; for Moore’s description (and it must be remembered that he was a first-rate judge) is clearly not applicable, as far as size of body and length of beak are concerned, to our present Jacobins. In 1795, judging from Bechstein, the breed had assumed its present character.

_Turbits._—It has generally been supposed by the older writers on pigeons, that the Turbit is the Cortbeck of Aldrovandi; but if this be the case, it is an extraordinary fact that the characteristic frill should not have been noticed. The beak, moreover, of the Cortbeck is described as closely resembling that of the Jacobin, which shows a change in the one or the other race. The Turbit, with its characteristic frill, and bearing its present name, is described by Willughby in 1677; and the bill is said to be like that of the bullfinch,—a good comparison, but now more strictly applicable to the beak of the Barb. The sub-breed called the Owl was well known in Moore’s time, in 1735.

_Tumblers._—Common Tumblers, as well as Ground Tumblers, perfect as far as tumbling is concerned, existed in India before the year 1600; and at this period diversified modes of flight, such as flying at night, the ascent to a great height, and manner of descent, seem to have been much attended to in India, as at the present time. Belon[40] in 1555 saw in Paphlagonia what he describes as “a very new thing, viz. pigeons which flew so high in the air that they were lost to view, but returned to their pigeon-house without separating.” This manner of flight is characteristic of our present Tumblers, but it is clear that Belon would have mentioned the act of tumbling if the pigeons described by him had tumbled. Tumblers were not known in Europe in 1600, as they are not mentioned by Aldrovandi, who discusses the flight of pigeons. They are briefly alluded to by Willughby, in 1687, as small pigeons “which show like footballs in the air.” The short-faced race did not exist at this period, as Willughby could not have overlooked birds so remarkable for their small size and short beaks. We can even trace some of the steps by which this race has been produced. Moore in 1735 enumerates correctly the chief points of excellence, but does not give any description of the several sub-breeds; and from this fact Mr. Eaton infers[41] that the Short-faced Tumbler had not then come to full perfection. Moore even speaks of the Jacobin as being the smallest pigeon. Thirty years afterwards, in 1765, in the Treatise dedicated to Mayor, short-faced Almond Tumblers are fully described, but the author, an excellent fancier, expressly states in his Preface (p. xiv.) that, “from great care and expense in breeding them, they have arrived to so great perfection and are so different from what they were 20 or 30 years past, that an old fancier would have condemned them for no other reason than because they are not like what used to be thought good when he was in the fancy before.” Hence it would appear that there was a rather sudden change in the character of the short-faced Tumbler at about this period; and there is reason to suspect that a dwarfed and half-monstrous bird, the parent-form of the several short-faced sub-breeds, then appeared. I suspect this because short-faced Tumblers are born with their beaks (ascertained by careful measurement) as short, proportionally with the size of their bodies, as in the adult bird; and in this respect they differ greatly from all other breeds, which slowly acquire during growth their various characteristic qualities.

Since the year 1765 there has been some change in one of the chief characters of the short-faced Tumbler, namely, in the length of the beak. Fanciers measure the “head and beak” from the tip of the beak to the front corner of the eyeball. About the year 1765 a “head and beak” was considered good,[42] which, measured in the usual manner, was 7/8 of an inch in length; now it ought not to exceed 5/8 of an inch; “it is however possible,” as Mr. Eaton candidly confesses,“for a bird to be considered as pleasant or neat even at 6/8 of an inch, but exceeding that length it must be looked upon as unworthy of attention.” Mr. Eaton states that he has never seen in the course of his life more than two or three birds with the “head and beak” not exceeding half an inch in length; “still I believe in the course of a few years that the head and beak will be shortened, and that half-inch birds will not be considered so great a curiosity as at the present time.” That Mr. Eaton’s opinion deserves attention cannot be doubted, considering his success in winning prizes at our exhibitions. Finally in regard to the Tumbler it may be concluded from the facts above given that it was originally introduced into Europe, probably first into England, from the East; and that it then resembled our common English