[75] Thus a male foetus showing reptilian characters in sexual ducts was exhibited by Shattock at the Pathological Society of London, February 19, 1895.
[76] J. Kohlbrugge, "Die Umgestaltung des Uterus der Affen nach den Geburt," _Zeitschrift für Morphologie_, bd. iv, p. 1, 1901.
[77] There are, however, no special nerve endings (Krause corpuscles), as was formerly supposed. The nerve endings in the genital region are the same as elsewhere. The difference lies in the abundance of superposed arboreal ramifications. See, e.g., Ed. Retterer, Art. "Ejaculation," Richet's _Dictionnaire de Physiologie_, vol. v.
[79] Sensations of pleasure without those of touch appear to be normal at the tip of the penis, as pointed out by Scripture, quoted in _Alienist and Neurologist_, January, 1898.
[80] See the previous volume of these _Studies_, "Sexual Selection in Man," p. 161.
[81] See, e.g., Ploss and Bartels, _Das Weib_, vol. i, beginning of chapter VI.
[82] Hyrtl states that the name _labia_ was first used by Haller in the middle of the eighteenth century in his _Elements of Physiology_, being adopted by him from the Greek poet Erotion, who gave these structures the very obvious name cheilea, lips. But this seems to be a mistake, for the seventeenth century anatomists certainly used the name "labia" for these parts.
[83] Bergh tentatively suggests, as regards the pubic hair, that its appearance may be due to the upright walk in man and the human position during coitus, the hair preventing irritation of the genitals from the sweat pouring down from the body and protecting the skin from direct friction in coitus. (In both these suggestions he was, however, long previously anticipated by Fabricius ab Aquapendente.) The fanciful suggestion of Louis Robinson that the pubic hair has developed in order to enable the human infant to cling securely to his mother is very poorly supported by facts, and has not met with acceptance. It may be mentioned that (as stated by Ploss and Bartels) the women of the Bismarck Archipelago, whose pubic hair is very abundant, use it as a kind of handkerchief on which to clean their hands.
[84] Routh and Heywood Smith have noted that the pubic hair tends to lose its curliness and become straight in women who masturbate. (_British Gynæcological Journal_, February, 1887, p. 505.)
[85] Schurig, _Muliebria_, p. 75. Plazzon in 1621 said that in Italian it had a popular name, _il besneegio_.
[86] Schurig brought together in his _Gynæcologia_ (pp. 2-4) various early opinions concerning the clitoris as the seat of voluptuous feeling.
[88] Adler, _Die Mangelhafte Geschlechtsempfindung des Weibes_, 1904, pp. 117-119.
[89] The voluptuous sensations caused by sexual contacts producing movements of the womb are probably normal and usual. They may even occur under circumstances unconnected with sexual emotion, and Mundé (_International Journal of Surgery_, March, 1893) mentions incidentally that in one case while titillating the cervix with a sound the woman very plainly showed voluptuous manifestations.
[90] Henle stated that fine hairs are frequently visible on the nymphæ; Stieda (_Zeitschrift für Morphologie_, 1902, p. 458) remarks that he has never been able to see them with the naked eye.
[91] R.L. Dickinson, "Hypertrophies of the Labia Minora and Their Significance," _American Gynæcologist_, September, 1902. It is perhaps noteworthy that Bergh found that in 302 cases in which the nymphæ were of unequal length, in all but 24 the left was longer.
[92] It may be remarked that Bergh believes that the nymphæ, and indeed the external genitals generally, are congenitally more strongly developed in libidinous persons, and at the same time in brunettes, while in public prostitutes this is not usually the case, which confirms the belief that exalted sexual sensibility does not usually lead to prostitution. He adds that prostitution, unless carried on for many years, has little effect on the shape of the external genitals.
[93] Schurig (_Muliebria_, 1729, Section II, cap. II) gives numerous quotations on this point; thus De Graaf wrote in his book on the sexual organs of women: "Tales protuberantiæ nymphæ appellantur ea propter quod aquis e vesica prosilientibus proxime adstare reperiantur, quandoquidem inter illas, tanquam duos parietes, urina magno impetu cum sibilo sæpe et absque labiorum irrigatione erumpit, vel quod sint castitatis præsides, aut sponsam primo intromittant."
[94] Havelock Ellis, "The Bladder as a Dynamometer," _American Journal of Dermatology_, May, 1902. If a woman who has never been pregnant, standing in the erect position before commencing the act of urination presses apart the labia minora with index and middle fingers the stream will be projected forward so as to fall usually at a considerable distance in front of a vertical line from the meatus; if when the act is half completed the fingers are removed, the labia close together and the stream, though maintained at a constant pressure, at once changes its character and direction.
[95] In poetry this term was employed by Plautus, _Pseudolus_, Act IV, Sc. 7. The Greek aidoion sometimes meant vagina and sometimes the external sexual parts; kolpos was used for the vagina alone.
[96] It is curious, however, that the European physicians of the seventeenth and even eighteenth centuries were doubtful of its value as a sign of virginity and considered it often absent.
[97] For a summary of the beliefs and practices of various peoples with regard to the hymen and virginity see Ploss and Bartels, _Das Weib_, vol. i, Chapter XVI.
II The Object of Detumescence--Erogenous Zones--The Lips--The Vascular Characters of Detumescence--Erectile Tissue--Erection in Woman--Mucous Emission in Women--Sexual Connection--The Human Mode of Intercourse--Normal Variations--The Motor Characters of Detumescence--Ejaculation--The Virile Reflex--The General Phenomena of Detumescence--The Circulatory and Respiratory Phenomena--Blood Pressure--Cardiac Disturbance--Glandular Activity--Distillatio--The Essentially Motor Character of Detumescence--Involuntary Muscular Irradiation to Bladder, etc.--Erotic Intoxication--Analogy of Sexual Detumescence and Vesical Tension--The Specifically Sexual Movements of Detumescence in Man--In Woman--The Spontaneous Movements of the Genital Canal in Woman--Their Function in Conception--Part Played by Active Movement of the Spermatozoa--The Artificial Injection of Semen--The Facial Expression During Detumescence--The Expression of Joy--The Occasional Serious Effects of Coitus.
We have seen what the object of detumescence is, and we have briefly considered the organs and structures which are chiefly concerned in the process. We have now to inquire what are the actual phenomena which take place during the act of detumescence.
Detumescence is normally linked closely to tumescence. Tumescence is the piling on of the fuel; detumescence is the leaping out of the devouring flame whence is lighted the torch of life to be handed on from generation to generation. The whole process is double and yet single; it is exactly analogous to that by which a pile is driven into the earth by the raising and then the letting go of a heavy weight which falls on to the head of the pile. In tumescence the organism is slowly wound up and force accumulated; in the act of detumescence the accumulated force is let go and by its liberation the sperm-bearing instrument is driven home. Courtship, as we commonly term the process of tumescence which takes place when a woman is first sexually approached by a man, is usually a highly prolonged process. But it is always necessary to remember that every repetition of the act of coitus, to be normally and effectively carried out on both sides, demands a similar double process; detumescence must be preceded by an abbreviated courtship.
This abbreviated courtship by which tumescence is secured or heightened in the repetition of acts of coitus which have become familiar, is mainly tactile.[98] Since the part of the man in coitus is more active and that of the woman more passive, the sexual sensitivity of the skin seems to be more pronounced in women. There are, moreover, regions of the surface of a woman's body where contact, when sympathetic, seems specially liable to arouse erotic excitement. Such erogenous zones are often specially marked in the breasts, occasionally in the palm of the hand, the nape of the neck, the lobule of the ear, the little finger; there is, indeed, perhaps no part of the surface of the body which may not, in some individuals at some time, become normally an erogenous zone. In hysteria the erotic excitability of these zones is sometimes very intense. The lips are, however, without doubt, the most persistently and poignantly sensitive region of the whole body outside the sphere of the sexual organs themselves. Hence the significance of the kiss as a preliminary of detumescence.[99] The importance of the lips as a normal erogenous zone is shown by the experiments of Gualino. He applied a thread, folded on itself several times, to the lips, thus stimulating them in a simple mechanical manner. Of 20 women, between the ages of 18 and 35, only 8 felt this as a merely mechanical operation, 4 felt a vaguely erotic element in the proceeding, 3 experienced a desire for coitus and in 5 there was actual sexual excitement with emission of mucus. Of 25 men, between the ages of 20 and 30, in 15 all sexual feeling was absent, in 7 erotic ideas were suggested with congestion of the sexual organs without erection, and in 3 there was the beginning of erection. It should be added that both the women and the men in whom this sexual reflex was more especially marked were of somewhat nervous temperament; in such persons erotic reactions of all kinds generally occur most easily. (Gualino, "Il Rifflesso Sessuale nell' eccitamento alle labbre," _Archivio di Psichiatria_, 1904, p. 341.)
As tumescence, under the influence of sensory stimulation, proceeds toward the climax when it gives place to detumescence, the physical phenomena become more and more acutely localized in the sexual organs. The process which was at first predominantly nervous and psychic now becomes more prominently vascular. The ancient sexual relationship of the skin asserts itself; there is marked surface congestion showing itself in various ways. The face tends to become red, and exactly the same phenomenon is taking place in the genital organs; "an erection," it has been said, "is a blushing of the penis." The difference is that in the genital organs this heightened vascularity has a definite and specific function to accomplish--the erection of the male organ which fits it to enter the female parts--and that consequently there has been developed in the penis that special kind of vascular mechanism, consisting of veins in connective tissue with unstriped muscular fibers, termed erectile tissue.[100] It is not only the man who is supplied with erectile tissue which in the process of tumescence becomes congested and swollen. The woman also, in the corresponding external genital region, is likewise supplied with erectile tissue now also charged with blood, and exhibits the same changes as have taken place in her partner, though less conspicuously visible. In the anthropoid apes, as the gorilla, the large clitoris and the nymphæ become prominent in sexual excitement, but the less development of the clitoris in women, together with the specifically human evolution of the mons veneris and larger lips, renders this sexual turgescence practically invisible, though it is perceptible to touch in an increased degree of spongy and elastic tension. The whole feminine genital canal, including the uterus, indeed, is richly supplied with blood-vessels, and is capable during sexual excitement of a very high degree of turgescence, a kind of erection.
The process of erection in woman is accompanied by the pouring out of fluid which copiously bathes all parts of the vulva around the entrance to the vagina. This is a bland, more or less odorless mucus which, under ordinary circumstances, slowly and imperceptibly suffuses the parts. When, however, the entrance to the vagina is exposed and extended, as during a gynæcological examination which occasionally produces sexual excitement, there may be seen a real ejaculation of the fluid which, as usually described, comes largely from the glands of Bartholin, situated at the mouth of the vagina. Under these circumstances it is sometimes described as being emitted in a jet which is thrown to a distance.[101] This mucous ejaculation was in former days regarded as analogous to the seminal ejaculation in man, and hence essential to conception. Although this belief was erroneous the fluid poured out in this manner whenever a high degree of tumescence is attained, and before the onset of detumescence, certainly performs an important function in lubricating the entrance to the genital canal and so facilitating the intromission of the male organ.[102] Menstruation has a similar influence in facilitating coitus, as Schurig long since pointed out.[103] A like process takes place during parturition when the same parts are being lubricated and stretched in preparation for the protrusion of the foetal head. The occurrence of the mucous flow in tumescence always indicates that that process is actively affecting the central sexual organs, and that voluptuous emotions are present.[104] The secretions of the genital canal and outlet in women are somewhat numerous. We have the odoriferous glands of sebaceous origin, and with them the prepuce of the clitoris which has been described as a kind of gigantic sebaceous follicle with the clitoris occupying its interior. (Hyrtl.) There is the secretion from the glands of Bartholin. There is again the vaginal secretion, opaque and albuminous, which appears to be alkaline when secreted, but becomes acid under the decomposing influence of bacteria, which are, however, harmless and not pathogenic. (Gow, _Obstetrical Society of London_, January 3, 1894.) There is, finally, the mucous uterine secretion, which is alkaline, and, being poured out during orgasm, is believed to protect the spermatozoa from destruction by the acid vaginal secretion.
The belief that the mucus poured out in women during sexual excitement is feminine semen and therefore essential to conception had many remarkable consequences and was widespread until the seventeenth century. Thus, in the chapter "De Modo coeundi et de regimine eorum qui coeunt" of _De Secretis Mulierum_, there is insistence on the importance of the proper mixture of the male semen with the female semen and of arranging that it shall not escape from the vagina. The woman must lie quiet for several hours at least, not rising even to urinate, and when she gets up, be very temperate in eating and drinking, and not run or jump, pretending that she has a headache. It was the belief in feminine semen which led some theologians to lay down that a woman might masturbate if she had not experienced orgasm in coitus. Schurig in his _Muliebria_ (1729, pp. 159, et seq.) discusses the opinions of old authors regarding the nature, source, and uses of the female genital secretions, and quotes authorities against the old view that it was female semen. In a subsequent work (_Syllepsilogia_, 1731, pp. 3, et seq.) he returns to the same question, quotes authors who accept a feminine semen, shows that Harvey denied it any significance, and himself decides against it. It has not seriously been brought forward since.
When erection is completed in both the man and the woman the conditions necessary for conjugation have at last been fulfilled. In all animals, even those most nearly allied to man, coitus is effected by the male approaching the female posteriorly. In man the normal method of male approach is anteriorly, face to face. Leonardo da Vinci, in a well-known drawing representing a sagittal section of a man and a woman connected in this position of so-called Venus obversa; has shown how well adapted the position is to the normal position of the organs in the human species.[105] Among monkeys, it is stated, congress is sometimes performed when the female is on all fours; at other times the male brings the female between his thighs when he is sitting, holding her with his forepaws. Froriep informed Lawrence that the male sometimes supported his feet on the female's calves. (Sir W. Lawrence, _Lectures on Physiology_, 1823, p. 186.) A summary of the methods of congress practiced by the various animals below mammals will be found in the article "Copulation" by H. de Varigny in Richet's _Dictionnaire de Physiologie_, vol. iv.
The anterior position in coitus, with the female partner lying supine, is so widespread throughout the world that it may fairly be termed the most typically human attitude in sexual congress. It is found represented in Egyptian graves at Benihassan, belonging to the Twelfth Dynasty; it is regarded by Mohammedans as the normal position, although other positions are permitted by the Prophet: "Your wives are your tillage: go in unto your tillage in what manner soever you will;" it is that adopted in Malacca; it appears, from Peruvian antiquities, to have been the position generally, though not exclusively, adopted in ancient Peru; it is found in many parts of Africa, and seems also to have been the most usual position among the American aborigines.
Various modifications of this position are, however, found. Thus, in some parts of the world, as among the Suahelis in Zanzibar, the male partner adopts the supine position. In Loango, according to Pechuel-Loesche, coitus is performed lying on the side. Sometimes, as on the west coast of Africa, the woman is supine and the man more or less erect; or, as among the Queenslanders (as described by Roth) the woman is supine and the man squats on his heels with her thighs clasping his flanks, while he raises her buttocks with his hands.
The position of coitus in which the man is supine is without doubt a natural and frequent variation of the specifically human obverse method of coitus. It was evidently familiar to the Romans. Ovid mentions it (_Ars Amatoria_, III, 777-8), recommending it to little women, and saying that Andromache was too tall to practice it with Hector. Aristophanes refers to it, and there are Greek epigrams in which women boast of their skill in riding their lovers. It has sometimes been viewed with a certain disfavor because it seems to confer a superiority on the woman. "Cursed be he," according to a Mohammedan saying, "who maketh woman heaven and man earth."
Of special interest is the wide prevalence of an attitude in coitus recalling that which prevails among quadrupeds. The frequency with which on the walls of Pompeii coitus is represented with the woman bending forward and her partner approaching her posteriorly has led to the belief that this attitude was formerly very common in Southern Italy. However that may be, it is certainly normal at the present day among various more or less primitive peoples in whom the vulva is often placed somewhat posteriorly. It is thus among the Soudanese, as also, in an altogether different part of the world, among the Eskimo Innuit and Koniags. The New Caledonians, according to Foley, cohabit in the quadrupedal manner, and so also the Papuans of New Guinea (Bongu), according to Vahness. The same custom is also found in Australia, where, however other postures are also adopted. In Europe the quadrupedal posture would seem to prevail among some of the South Slavs, notably the Dalmatians. (The different methods of coitus practiced by the South Slavs are described in Kryptadia vol. vi, pp. 220, et seq.)
This method of coitus was recommended by Lucretius (lib. iv) and also advised by Paulus Æginetus as favorable to conception. (The opinions of various early physicians are quoted by Schurig, _Spermatologia_, 1720, pp. 232, et seq.). It seems to be a position that is not infrequently agreeable to women, a fact which may be brought into connection with the remarks of Adler already quoted (p. 131) concerning the comparative lack of adjustment of the feminine organs to the obverse position. It is noteworthy that in the days of witchcraft hysterical women constantly believed that they had had intercourse with the Devil in this manner. This circumstance, indeed, probably aided in the very marked disfavor in which coitus _a posteriori_ fell after the decay of classic influences. The mediæval physicians described it as _mos diabolicus_ and mistakenly supposed that it produced abortion (Hyrtl, op. cit., vol. ii, p. 87). The theologians, needless to say, were opposed to the _mos diabolicus_, and already in the Anglo-Saxon Penitential of Theodore, at the end of the seventh century, 40 days' penance is prescribed for this method of coitus.
From the frequency with which they have been adopted by various peoples as national customs, most of the postures in coitus here referred to must be said to come within the normal range of variation. It is a mistake to regard them as vicious perversions.
Up to the point to which we have so far considered it, the process of detumescence has been mainly nervous and vascular in character; it has, in fact, been but the more acute stage of a process which has been going on throughout tumescence. But now we reach the point at which a new element comes in: muscular action. With the onset of muscular action, which is mainly involuntary, even when it affects the voluntary muscles, detumescence proper begins to take place. Henceforward purposeful psychic action, except by an effort, is virtually abolished. The individual, as a separate person, tends to disappear. He has become one with another person, as nearly one as the conditions of existence ever permit; he and she are now merely an instrument in the hands of a higher power--by whatever name we may choose to call that Power--which is using them for an end not themselves.
The decisive moment in the production of the instinctive and involuntary orgasm occurs when, under the influence of the stimulus applied to the penis by friction with the vagina, the tension of the seminal fluid poured into the urethra arouses the ejaculatory center in the spinal cord and the bulbo-cavernosus muscle surrounding the urethra responsively contracts in rhythmic spasms. Then it is that ejaculation occurs.[106] "The circulation quickens, the arteries beat strongly," wrote Roubaud in a description of the physical state during coitus which may almost be termed classic; "the venous blood, arrested by muscular contraction, increases the general heat, and this stagnation, more pronounced in the brain by the contraction of the muscles of the neck and the throwing of the head backward, causes a momentary cerebral congestion, during which intelligence is lost and the faculties abolished. The eyes, violently injected, become haggard, and the look uncertain, or, in the majority of cases, the eyes are closed spasmodically to avoid the contact of the light. The respiration is hurried, sometimes interrupted, and may be suspended by the spasmodic contraction of the larynx, and the air, for a time compressed, is at last emitted in broken and meaningless words. The congested nervous centers only communicate confused sensations and volitions; mobility and sensation show extreme disorder; the limbs are seized by convulsions and sometimes by cramps, or are thrown wildly about or become stiff like iron bars. The jaws, tightly pressed, grind the teeth, and in some persons the delirium is carried so far that they bite to bleeding the shoulders their companions have imprudently abandoned to them. This frantic state of epilepsy lasts but a short time, but it suffices to exhaust the forces of the organism, especially in man. It is, I believe, Galen, who said: 'Omne animal post coitum triste præter mulierem gallumque.'"[107] Most of the elements that make up this typical picture of the state of coitus are not absolutely essential to that state, but they all come within the normal range of variation. There can be no doubt that this range is considerable. There would appear to be not only individual, but also racial, differences; there is a remarkable passage in Vatsyayana's _Kama Sutra_ describing the varying behavior of the women of different races in India under the stress of sexual excitement--Dravidian women with difficulty attaining erethism, women of the Punjaub fond of being caressed with the tongue, women of Oude with impetuous desire and profuse flow of mucus, etc.--and it is highly probable, Ploss and Bartels remark, that these characterizations are founded on exact observations.[108] The various phenomena included in Roubaud's description of the condition during coitus may all be directly or indirectly reduced to two groups: the first circulatory and respiratory, the second motor. It is necessary to consider both these aspects of the process of detumescence in somewhat greater detail, although while it is most convenient to discuss them separately, it must be borne in mind that they are not really separable; the circulatory phenomena are in large measure a by-product of the involuntary motor process.
With the approach of detumescence the respiration becomes shallow, rapid, and to some extent arrested. This characteristic of the breathing during sexual excitement is well recognized; so that in, for instance, the _Arabian Nights_, it is commonly noted of women when gazing at beautiful youths whose love they desired, that they ceased breathing.[109] It may be added that exactly the same tendency to superficial and arrested respiration takes place whenever there is any intense mental concentration, as in severe intellectual work.[110] The arrest of respiration tends to render the blood venous, and thus aids in stimulating the vasomotor centers, raising the blood-pressure in the body generally, and especially in the erectile tissues. High blood-pressure is one of the most marked features of the state of detumescence. The heart beats are stronger and quicker, the surface arteries are more visible, the conjunctivæ become red. The precise degree of blood-pressure attained during coitus has been most accurately ascertained in the dog. In Bechterew's laboratory in St. Petersburg a manometer was introduced into the central end of the carotid artery of a bitch; a male dog was then introduced, and during coitus observations were made on the blood-pressure at the peripheral and central ends of the artery. It was found that there was a great general elevation of blood-pressure, intense hyperæmia of the brain, rapid alternations, during the act, of vasoconstriction and vasodilatation of the brain, with increase and diminution of the general arterial tension in relation with the various phases of the act, the greatest cerebral vasodilatation and hyperæmia coinciding with the moment following the intromission of the penis; the end of the act is followed by a considerable fall in the blood-pressure.[111] I am not acquainted with any precise observations on the blood-pressure in human subjects during detumescence, and there are obvious difficulties in the way of such observations. It is probable, however, that the conditions found would be substantially the same. This is indicated, so far as the very marked increase of blood-pressure is concerned, by some observations made by Vaschide and Vurpas with the sphygmanometer on a lady under the influence of sexual excitement. In this case there was a relationship of sympathy and friendly tenderness between the experimenter and the subject, Madame X, aged 25. Experimenter and subject talked sympathetically, and finally, we are told, while the latter still had her hands in the sphygmanometer, the former almost made a declaration of love. Madame X was greatly impressed, and afterward admitted that her emotions had been genuine and strong. The blood-pressure, which was in this subject habitually 65 millimeters, rose to 150 and even 160, indicating a very high pressure, which rarely occurs; at the same time Madame X looked very emotional and troubled.[112] Some authorities are of opinion that irregularities in the accomplishment of the sexual act are specially liable to cause disturbances in the circulation. Thus Kisch, of Prague, refers to the case of a couple practising coitus interruptus--the husband withdrawing before ejaculation--in which the wife, a vigorous woman, became liable after some years to attacks termed by Kisch _neurasthenia cordis vasomotoria_, in which there was at daily or longer intervals palpitation, with feelings of anxiety, headache, dizziness, muscular weakness and tendency to faint. He regards coitus as a cause of various heart troubles in women: (1) Attacks of tachycardia in very excitable and sexually inclined women; (2) attacks of tachycardia with dyspnoea in young women, with vaginismus; (3) cardiac symptoms with lowered vascular tone in women who for a long time have practised coitus interruptus without complete sexual gratification (Kisch, "Herzbeschwerden der Frauen verursacht durch den Cohabitationsact," _Münchener Medizinisches Wochenschrift_, 1897, p. 617). In this connection, also, reference may probably be made to those attacks of anxiety which Freud associates with psychic sexual lesions of an emotional character.
Associated with this vascular activity in detumescence we find a general tendency to glandular activity. Various secretions are formed abundantly. Perspiration is copious, and the ancient relationship between the cutaneous and sexual systems seems to evoke a general activity of the skin and its odoriferous secretions. Salivation, which also occurs, is very conspicuous in many lower animals, as for instance in the donkey, notably the female, who just before coitus stands with mouth open, jaws moving, and saliva dribbling. In men, corresponding to the more copious secretion in women, there is, during the latter stages of tumescence, a slight secretion of mucus--Fürbringer's _urethrorrhoea ex libidine_--which appears in drops at the urethral orifice. It comes from the small glands of Littré and Cowper which open into the urethra. This phenomenon was well known to the old theologians, who called it _distillatio_, and realized its significance as at once distinct from semen and an indication that the mind was dwelling on voluptuous images; it was also known in classic times[113]; more recently it has often been confused with semen and has thus sometimes caused needless anxiety to nervous persons. There is also an increased secretion of urine, and it is probable that if the viscera were more accessible to observation we might be able to demonstrate that the glands throughout the body share in this increased activity.
The phenomena of detumescence culminate, however, and have their most obvious manifestation in motor activity. The genital act, as Vaschide and Vurpas remark, consists essentially in "a more and more marked tension of the motor state which, reaching its maximum, presents a short tonic phase, followed by a clonic phase, and terminates in a period of adynamia and repose." This motor activity is of the essence of the impulse of detumescence, because without it the sperm cells could not be brought into the neighborhood of the germ cell and be propelled into the organic nest which is assigned for their conjunction and incubation.